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低温 ていおん生物 せいぶつ学 がくでした
teionseibutsugakudeshita

敬
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Noun
, 
Desu Copula

The polite past form of 低温生物学

1. Low-temperature biology was.; It was low-temperature biology.

Past/Perfective
, 
Polite

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低温生物学でした

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