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行動 こうどう生態 せいたい学 がくでした
kōdō seita igakudeshita

敬
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Noun
, 
Desu Copula

The polite past form of 行動生態学

1. Behavioral ecology was. ; ; It was behavioral ecology.

Past/Perfective
, 
Polite

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行動生態学でした

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